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Genus CAMPONOTUS Mayr

     Camponotus is a huge, worldwide genus of bewilderingcomplexity. Many species are quite variable and most of the variants have been named, resulting in a welter of inadequately described, poorly understood forms whose relationships are often
obscured by superficial differences which have little or no taxonomic value. This genus is an excellent example of taxonomic chaos.
     Fortunately, the forms in the United States are, for the most part, in a relatively stable taxonomic condition. The group of species comprising the subgenus Myrmentoma, was revised by Snelling (1988). We have but two species of Myrmentoma in the desert and their separation presents no problems.
     The remaining species are fairly easy to separate. The key includes C. (Colobopsis) papago Creighton, an arboreal species not known to occur in our area. Since it has been taken in Arizona near the Colorado River it seemed well to include it as a potential member of our fauna.

KEY TO SPECIES OF CAMPONOTUS


 
Workers
 
1 Head of major, in profile, evenly arched from clypeal apex to vertex, lower area not sharply contrastingly sculptured

2
-- Head of major, in profile, obliquely truncate below, truncation marginate and involving most of clypeus and adjacent side of face, truncated portion vermiculate-rugose or reticulose, sharply contrasting with sculpture elsewhere on head (subg. Colobopsis)


papago Creighton*
2 (1) Lower margin of median lobe of clypeus thickened, either transverse or slightly convex and irregularly eroded or subdentate (Figs. 210, 212, 218) (subg. Tanaemyrmex)


3
-- Lower margin of median lobe of clypeus thin, weakly convex and semicircularly depressed and incised in middle (Figs. 214, 216) (subg. Myrmentoma)

7
3 (2) Malar area without erect hairs or with no more than two or three; lower margin of clypeus slightly convex, irregular; scape without erect hairs except several tip


4
-- Malar area with 15+ erect hairs; median lobe of clypeus set off on each side by distinct angulation, its ventral margin transverse, even; scape with scattered erect hairs (Fig. 210)

festinatus (Buckley)
4(3) Metatibia usually not contrastingly darker than femur; median carina of clypeus not terminating in blunt tooth; pronotum usually with 10 or more erect hairs and node of petiole with five or six on each side; scape usually flattened at base




5

--�������� Metatibia blackish, contrasting with femur; median carina of clypeus ending in blunt tooth; pronotum with fewer than ten, often no, erect hairs and scale of petiole with no more than two or three, often no, erect hairs on each side; scape not flattened at base (Fig. 212)


ocreatus Emery
5(4) Base of scape flattened, at least slightly broader at base than immediately beyond; pubescence of T2 usually shorter than distance between individual hairs



6
--
Scape narrowest at base, weakly flattened (Fig. 220); discal pubescence of T2 longer than distance between individual hairs

vicinus Mayr
6(5) Margins of head of major, in frontal view, distinctly convergent toward mandibular bases; scape with small lateral lobule at base; apex of scape of major exceeding vertex corner by at least its apical breadth (Fig. 219)





semitestaceus Snelling
-- Head of major quadrate, margins strongly convergent immediately above mandibular base; scape base broad, flattened, not lobulate; apex of scape of major exceeding vertex corner, if at all, by less than its apical breadth (Fig. 218)



sansabeanus (Buckley)
7(2) Metanotum, in profile, distinctly depressed below adjacent mesonotum and propodeum; outer face of mandible with round punctures, interspaces often polished; clypeus of major flattened and shiny (Figs. 213, 214)



hyatti
Emery
-- Metanotum, in profile, little, if any, depressed; outer face of mandible with elongate punctures, interspaces dull; clypeus of major convex, dull (Figs. 215, 216)

sayi
Emery
 
females

1 Head, in profile, evenly arched from lower clypeal margin to vertex; lower portions neither vermiculate-rugose nor reticulose .

2
-- Head, in profile, obliquely truncate below, truncation marginate and involving most of clypeus and adjacent sides of face and vermiculate-rugose or reticulose (subg. Colobopsis)

papago Creighton*
2(1) Ventral margin of median lobe of clypeus thickened, either transverse or slightly convex and irregularly eroded or subdentate (subg. Tanaemyrmex)


3
-- Ventral margin of median lobe of clypeus thin, weakly convex and semicircularly depressed and incised in middle (subg. Myrmentoma)

7
3(2) Malar area without erect hairs, or with fewer than three; ventral margin of median lobe of clypeus slightly convex and irregularly eroded

4
-- Malar area with 6-15 erect hairs; median lobe of clypeus angulate at sides, ventral margin transverse; scape with scattered erect hairs

festinatus (Buckley)
4(3) Median carina of clypeus, if present, not terminating as dentiform process; base of scape flattened; summit of petiole node usually entire; tibiae no darker than femora


5
-- Median carina of clypeus terminating in dentiform process; scape cylindrical at base; petiolar node incised at summit; tibiae darker than femora

ocreatus Emery
5(4) Scape surpassing vertex corner by more than length of A2; front of head contiguously micropunctate, subgranulose, or finely and closely tessellate, punctures sparse, fine and obscure; mesosomal side slightly to moderately shiny, surface distinctly tessellate




6
-- Scape surpassing vertex corner by less than length of A2; front of head moderately shiny, with sparse sharply defined small punctures and scattered coarse punctures; mesosomal side shiny, surface weakly tessellate in most areas


sansabeanus (Buckley)
6(5) Scape base flattened, with small lateral lobule; front of head finely, densely tessellate, usually not appearing granulose


semitestaceus Snelling
--
Scape base flattened, without lobule; front of head contiguously micropunctate, subgranulose

vicinus Mayr
7(2) Head and mesosoma reddish; ocelli large, IOD about 3 x OD, OOD no more than 4 x OD

sayi Emery
-- Head and mesosoma brownish; ocelli small, IOD at least 4.5 x OD, OOD about 6.5 x OD
hyatti Emery


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• • •
Date of this version 8, Nov. 2003
• • •
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